The 335th Regiment was originally constituted as the 335th Infantry in the National Army on 5 August 1917 and assigned to the 84th Division. The Regiment, along with the rest of the Division, was broken up after it arrived in France and its troops used as replacement cadres for units already in combat, which is why the Regiment’s World War I campaign streamer has no inscription for specific battles or campaigns in which the unit fought.
After being demobilized in 1919, the Regiment was reformed with the same in the Organized Reserves and once again assigned to the 84th Division, which became the 84th Airborne Division in 1942. It fought in three campaigns (Rhineland, Ardennes-Alsace, Central Europe) during World War II and was awarded a French Croix de Guerre with Palm for its courageous service during combat in Belgium. On 23 January 1946, it was inactivated at Camp Kilmer, New Jersey, and subsequently redesignated as the 335th Parachute Infantry.
The 335th Regiment has been allotted to the Reserve ever since its reactivation in 1947. It was briefly designated as the 335th Airborne Infantry (March 1951 to March 1952) before being reorganized and redesignated as the 335th Infantry yet again, but this time was relieved for assignment to the 84th Airborne Division and assigned to the 85th Division.
From 1959 until April 2007, the Regiment was organized so that it or its Battalion were designated as elements of the 85th Division, a training unit variously designated as Training, Exercise, or Training Support. The Regiment was relieved from assignment to the 85th Division when the 85th was redesignated as the 85th Support Command, U.S. Army Reserve in 2007; instead, the Regiment’s Battalions were assigned to the 157th Infantry Brigade, a subordinate unit of the 85th Support Command that also conducts training.
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More commonly known as a unit crest, the 335th Regiment Distinctive Unit Insignia (DUI) was originally approved for the 33th Infantry Regiment on 29 June 1926. It was redesignated for the 335th Airborne Infantry Regiment on 31 March 1952 before being redesignated for the 335th Regiment on 8 August 1960. On 3 April 1970, the insignia was amended to add the accent mark to the motto. Full guidance on how the DUI is to be worn can found in DA Pamphlet 670-1,
Section 21-22, "Distinctive unit insignia."A dark blue shield stand signifies the organization’s Infantry origins and assignment to Infantry units for deployment into combat. The three fleurs-de-lis and two gold horizontal bars with the blue stripe between them are adapted from the coat of arms of Le Mans, with the fleurs-de-lis denoting the World War I campaigns the Regiment fought in.
In the lower half of the insignia are five ermines; combined with the three fleurs-de-lis and the three made up of the two gold bars with blue stripe, it denotes the numerical designation of the organization. The Institute of Heraldry notes that the motto ‘A FIN translates into English as “To The End” but does not name the original language.